Joyce, F. H., R. A. Zahawi, and K. D. Holl. (2025) Lower-intensity restoration interventions drive greater seedling establishment for later-successional tree species. Journal of Applied Ecology https://doi.org/10.1111/1365-2664.70116 

Schubert, Spencer C., Rakan A. Zahawi, Federico Oviedo-Brenes, Juan Abel Rosales, and Karen D. Holl. (2025).  Active Restoration Increases Tree Species Richness and Recruitment of Large-Seeded Taxa after 16–18 years. Ecological Applications 35(1): e3053. https://doi.org/10.1002/eap.3053

Lackmann J.A., Fernandez Barrancos E.P., Zahawi R.A., Aldrich-Wolfe L. (2025) Restoration plantations accelerate recovery of fungal communities of coarse woody debris in southern Costa Rica. 58:e03487

Werden, L. K., Cole, R. J., Schönhofer, K., Holl, K. D., Zahawi, R. A., Averill, C., Schweizer, D., Calvo-Alvarado, J. C., Hamilton, D., Joyce, F. H., San-José, M., Hofhansl, F., Briggs, L., Rodríguez, D., Tingle, J. W., Chiriboga, F., Broadbent, E. N., Quirós-Cedeño, G. J., & Crowther, T. W. (2024). Assessing innovations for upscaling forest landscape restoration. One Earth, 7(9), 1515-1525. https://doi.org/10.1016/j.oneear.2024.07.011

Cole, R. J., L. K. Werden, F. C. Arroyo, K. M. Quirós, G. Q. Cedeño and T. W. Crowther (2024). Forest restoration in practice across Latin America. Biological Conservation 294: 11.https://doi.org/10.1016/j.biocon.2024.110608

Holl KD, Reid JL, Zahawi RA, De Siqueira LP, Brancalion PHS (2024) A call for practical spatially patterned forest restoration methods. Restoration Ecology 32:e14266 https://doi.org/10.1111/rec.14266 

Quirós Cedeño GJ, Joyce FH, Zúñiga Amador MA, Holl KD (2025) Differing Agents of Physical Damage to Artificial Seedlings in Remnant and Restored Forests. Biotropica 57:e13421. https://doi.org/10.1111/btp.13421

Joyce F. H., Rosales, J. A., Holl, K.D., Zahawi, R.A., Bui, A. and Reid, J. L. (2024). Active restoration accelerates recovery of tropical forest bird assemblages over two decades. Biological Conservation, https://doi.org/10.1016/j.biocon.2024.110593

Joyce, F. H., Ramos, B. M., Zahawi, R. A., & Holl, K. D. (2024). Vertebrate seed predation can limit recruitment of later-successional species in tropical forest restoration. Biotropica, 56, e13381. https://doi.org/10.1111/btp.13381

Schweizer, D., Cole, R.J., Werden, L.K., Cedeño, G.Q., Rodriguez, D., Navarro, K., Esquivel, J.M., Max, S., Chiriboga, F.E., Zahawi, R.A., Holl, K.D. and Crowther, T.W. (2024), Review and assessment of smartphone apps for forest restoration monitoring. Restor Ecol, 32: e14136. https://doi.org/10.1111/rec.14136

Tucker, N.I.J., Elliott, S., Holl, K.D., Zahawi, R.A. (2023). Restoring Tropical Forests: Lessons Learned from Case Studies on Three Continents. In: Florentine, S., Gibson-Roy, P., Dixon, K.W., Broadhurst, L. (eds) Ecological Restoration. Springer, Cham. https://doi.org/10.1007/978-3-031-25412-3_3

Kulikowski Andy J., Zahawi Rakan A., Werden Leland K., Zhu Kai and Holl Karen D. (2023) Restoration interventions mediate tropical tree recruitment dynamics over timePhil. Trans. R. Soc. B37820210077http://doi.org/10.1098/rstb.2021.0077

San-José, M., Werden, L.K., Joyce, F.H. et al. (2022) Effects of landscape structure on restoration success in tropical premontane forest. Sci Rep 12, 13452. https://doi.org/10.1038/s41598-022-16542-3

Kulikowski, A.J., II, Zahawi, R.A. and Holl, K.D. (2022), Effects of insect herbivory on seedling mortality in restored and remnant tropical forest. Restor Ecol, 30: e13467. https://doi.org/10.1111/rec.13467

Reid, J. L., R. A. Zahawi, D. A. Zárrate-Chary, J. A. Rosales, K. D. Holl, and U. Kormann. (2021). Multi-scale habitat selection of key frugivores predicts large-seeded tree recruitment in tropical forest restoration. Ecosphere 12(12):e03868. 10.1002/ecs2.3868

Coffee pulp accelerates early tropical forest succession on old fields. RJ Cole, RA Zahawi - Ecological Solutions and Evidence, 2021

Werden LK, Holl KD, Chaves-Fallas JM, Oviedo-Brenes F, Rosales JA, Zahawi RA. (2021) Degree of intervention affects interannual and within-plot heterogeneity of seed arrival in tropical forest restoration. J Appl Ecol. 58: 1693–1704. https://doi.org/10.1111/1365-2664.13907

Zahawi, R.A., Werden, L.K., San-José, M., Rosales, J.A., Flores, J. and Holl, K.D. (2021), Proximity and abundance of mother trees affects recruitment patterns in a long-term tropical forest restoration study. Ecography, 44: 1826-1837. https://doi.org/10.1111/ecog.05907

Assisted restoration interventions drive functional recovery of tropical wet forest tree communities. LK Werden, S Zarges, KD Holl, CL Oliver… - Frontiers in Forests and Global Change, 2022.

The landscape genetic signature of pollination by trapliners: evidence from the tropical herb, Heliconia tortuosa. F Torres-Vanegas, AS Hadley, UG Kormann, FA Jones… - Frontiers in Genetics, 2019

Applied nucleation facilitates tropical forest recovery: Lessons learned from a 15‐year study. KD Holl, JL Reid, RJ Cole, F Oviedo‐Brenes… - Journal of Applied Ecology, 2020

Ant–scale mutualism increases scale infestation, decreases folivory, and disrupts biological control in restored tropical forests. AJ Kulikowski - Biotropica, 2020

Leaf litter arthropod responses to tropical forest restoration. RJ Cole, KD Holl, RA Zahawi, P Wickey, AR Townsend - Ecology and Evolution, 2016

Seed dispersal limitations shift over time in tropical forest restoration. JL Reid, KD Holl, RA Zahawi - Ecological Applications, 2015

Testing applied nucleation as a strategy to facilitate tropical forest recovery. RA Zahawi, KD Holl, RJ Cole, JL Reid - Journal of Applied Ecology, 2013

Seed rain under tree islands planted to restore degraded lands in a tropical agricultural landscape. RJ Cole, KD Holl, RA Zahawi - Ecological Applications, 2010

Direct seeding of late-successional trees to restore tropical montane forest. RJ Cole, KD Holl, CL Keene, RA Zahawi - Forest Ecology and Management, 2011